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Question: Lack's model for clutch size evolution predicts the optimal number of offspring in a clutch as those which produce the most offspring. However, experiments often demonstrate that organisms do not produce the predicted optimal clutch size because:
his assumption that offspring receive equal parental investment is often not met.
his model assumes that there are no carry-over costs among reproductive events, yet early reproductive investment can affect future reproductive success.
his model uses offspring survival as a measure of fitness which may be less important in the long run than, for example, the quality of those offspring.
tests of his model often use approaches like adding eggs to nest, which may underestimate the true cost of increases reproductive investment.
All of the above.
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